Reference: Moretti M., Dias A.T.C., de Bello F., Altermatt F., Chown S.L., Azcarate F.M., Bell J.R., Fournier B., Hedde M., Hortal J., Ibanez S., Ockinger E., Sousa J.P., Ellers J. & Berg M.P. 2017. Handbook of protocols for standardized measurement of terrestrial invertebrate functional traits. Functional Ecology 31(3):558-567.